我是个菜鸟，稍微分享一下我的看法：Those fights are over mates,①which is ②what Dr Arboursuggests was also going on with ankylosaurs.①处作非限制性定语从句，which确实指代前面整个主句，定语从句谓语动词为紧跟在后面的is。②处为which is后的表语从句，但是结构还可以再划分：（what Dr Arbour suggests）was also going onwith ankylosaurs括号处又是主语从句，所以was是②处表语从句的谓语动词，suggests是主语从句的谓语动词。怎么翻译一下句子就是：这些争斗是为了（争夺）配偶（主句），Dr Arbour的言下之意是这对甲龙也同样适用。