关于代词that及平行省略

各位好,

Heartless human beings are like wild animals, which rip their own kin in the slightest of need and that too without a sign of remorse.

①这里that语义上应该是指代human beings对吗?可是不应该用those吗?

②就算是用了those,too的作用又是什么呢?中间省略了什么呢?

感谢各位的解答!

如题
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